Malacologia

HARVARD UNIVERSITY Library of the Museum of Comparative ZoologyNO. 1-2/OL. 34, 1992 MALACOLOGIA nternational Journal of Malacology Revista Internacional de Malacologia Journal International de Malacologie Internationale Malakologische ZeitschriftPublication dates Vol. No. 1-228, 19 January 1988 Vol. No. 129, 28 June 1988 Vol. No. 229, 16 Dec. 1988 Vol. 30, No. 1-2 1 Aug. 1989 Vol. 31, No. 1 29 Dec. 1989 Vol. 31, No. 2 28 May 1990 Vol. 32, No. 2 7 June 1991 Vol. 33, No. 1-2 6 Sep. 199119921-2'OL 34, NO. MCZ LIBRARY SEP 51 1992 HARVARD UNIVERSITY MALACOLOGIA Malacologyiternational Journal of MalacologiaRevista Internacional de Journal International de Malacologie ZeitschriftMalakologischeInternationaleMALACOLOGIA Editor-in-Chief: GEORGE M. DAVIS Editorial and Subscription Offices: Department of Malacology The Academy of Natural Sciences of Philadelphia 1900 Benjamin Franklin Parkway Philadelphia, Pennsylvania 19103-1195, U.S.A. Co-Editors: EUGENE COAN CAROL JONES California Academy of Sciences Denver, CO San Francisco, CA Assistant Managing Editor: CARYL HESTERMAN Associate Editors: JOHN B. BURCH ANNE GISMANN University of Michigan Maadi Ann Arbor Egypt MALACOLOGIA is published by the INSTITUTE OF MALACOLOGY, the Sponsor Members of which (also serving editors) are:as KENNETH J. BOSS JAMES NYBAKKEN, President Museum of Comparative Zoology Moss Landing Marine Laboratory Cambridge, Massachusetts California JOHN BURCH, President-Eiect CLYDE F. E. ROPER Smithsonian Institution MELBOURNE R. CARRIKER Washington, D.C. University of Delaware, Lewes W. D. RUSSELL-HUNTERGEORGE M. DAVIS Syracuse University, New York Secretaryand Treasurer SHI-KUEI WU CAROLE S. HICKMAN, Vice-President University of Colorado Museum, Boulder University of California, Berkeley Participating Members EDMUND GITTENBERGER JACKIE L VAN GOETHEM Secretary, UNITAS MALACOLOGICA Treasurer, UNITAS MALACOLOGICA Rijksmuseum van Natuurlijke Koninklijk Belgisch Instituut Historie voor Natuurwetenschappen Leiden, Netherlands Brüssel, Belgium Emeritus Members J. FRANCIS ALLEN, Emérita ROBERT ROBERTSON Environmental Protection Agency The Academy of Natural Sciences Washington, D.C. Philadelphia, Pennsylvania ELMER G. BERRY, NORMAN F. SOHL Germantown, Maryland U.S. Geological Survey Reston, Virginia Copyright © 1992 by the Institute of Malacology1992 EDITORIAL BOARD E. GITTENBERGERJ. A. ALLEN Biological Station Rijksmuseum van Natuurlijke HistorieMarine Millport, United Kingdom Leiden, Netherlands F. GIUSTI R. BIELER Université di Siena, Italy Field Museum Chicago, U.S.A. A. N. GOLIKOV Zoological Institute E. E. BINDER Leningrad, U.S.S.R. Muséum d'Histoire Naturelle S. J. GOULD Genève, Switzerland Harvard University Cambridge, Mass., U.S.A. A. J. CAIN University of Liverpool V.A. GROSSU United Kingdom Universitatea Bucuresti Romania P. CALOW T HABE University of Sheffield Tokai University United Kingdom Shimizu, Japan G. CARTER R. HANLONJ. University of North Carolina Marine Biomedical Institute Chapel Hill, U.S.A. Galveston, Texas, U.S.A. J. A. HENDRICKSON, Jr. R. COWIE Academy of Natural Sciences Bishop Museum Philadelphia, PA, U.S.A. Honolulu, HL, U.S.A. D. M. HILLIS University of TexasCLARKE, Jr.A. H. Austin, U.S.A.Portland, Texas, U.S.A. K. E. HOAGLAND CLARKEB. CollectionsAssociation of Systematics University of Nottingham Washington, DC, U.S.A. United Kingdom B. HUBENDICK Naturhistoriska Museet R. DILLON Göteborg, Sweden College of Charleston SC, U.S.A. S. HUNT Lancashire DUNCAN United KingdomC. J. University of Liverpool R. JANSSEN United Kingdom Forschungsinstitut Senckenberg, Frankfurtam Main, Germany D. J. EERNISSE R. N. KILBURN University of Michigan Natal MuseumAnn Arbor, U.S.A. Pietermaritzburg, South Africa V. FRETTER M. A. KLAPPENBACH University of Reading Museo Nacional de Historia Natural United Kingdom Montevideo, UruguayD. G. REIDJ. KNUDSEN History MuseumZoologisk Institut & Museum The Natural London, United KingdomKobenhavn, Denmark A. J. KOHN N. W. RUNHAM University of Washington University College of North Wales U.S.A.Seattle, Bangor, United Kingdom A. LUCAS S. G. SEGERSTRÂLE Faculté des Sciences Institute of Marine Research Brest, France Helsinki, Finland C. MEIER-BROOK STAÑCZYKOWSKAA. InstitutTropenmedizinisches Siedlce, Poland Tübingen, Germany F. STARMÜHLNERH. . MIENIS Institut der UniversitätZoologischesHebrew University ofJerusalem AustriaWien,Israel Y. STAROBOGATOVI.MORTONJ. E. Zoological InstituteThe University Leningrad, U.S.S.R.Auckland, New Zealand W. STREIFFMURRAY, Jr.J. J. Université de Caenof VirginiaUniversity FranceCharlottesville, U.S.A. NATARAJAN J. STUARDOR. ChileBiological Station Universidad deMarine Novo, India ValparaisoPorto J. 0KLAND S. TILLIER University of Oslo NaturelleMuséum National d'Histoire Norway Paris, France T. OKUTANI TURNERR. D. University of Fisheries UniversityHarvard Tokyo, Japan Cambridge, Mass., U.S.A. W. L. PARAENSE J.A.M. van den BIGGELAAR Instituto Oswaldo Cruz, Rio de Janeiro of UtrechtUniversity Brazil The Netherlands J. J. PARODIZ J. A. van EEDEN Carnegie Museum Potchefstroom University Pittsburgh, U.S.A. South Africa J. P. POINTER N. H. VERDONK Ecole Pratique des Hautes Etudes Rijksuniversiteit Perpignan Cedex, France Utrecht, Netherlands W. F. PONDER B. R. WILSONAustralian Museum and Land ManagementDept. ConservationSydney Netherlands, Western Australia R. D. PURCHON H. ZEISSLERTechnologyChelsea College of Science & Leipzig, GermanyKingdomLondon, United ZILCHY. A.Ql Z. Forschungsinstitut SenckenbergAcademia Sínica Frankfurtam Main, GermanyQingdao, People's Republic of China